/* =============================================================
   Quercus robur (QroDB) — 夏栎主题样式
   设计语言参照 WatermelonOmics：Hero + 统计条 + 模块卡 + 数据面板
   配色替换为夏栎：树皮棕 / 橡实金 / 叶绿
   ============================================================= */

:root {
  /* 树皮棕 — 品牌主色 */
  --oak-900: #2b1d0e;
  --oak-800: #4a3319;
  --oak-700: #6b4f2a;
  --oak-600: #8a6a3c;
  --oak-500: #a98a55;
  --oak-400: #c6ac82;
  --oak-300: #dfcdae;
  --oak-200: #efe4d0;
  --oak-100: #f7f0e3;
  --oak-50:  #fbf7ef;

  /* 叶绿 — 辅助主色 */
  --leaf-700: #23512c;
  --leaf-600: #2f6b3a;
  --leaf-500: #3f8a4c;
  --leaf-400: #63a86e;
  --leaf-300: #93c69b;
  --leaf-200: #c6e3ca;
  --leaf-100: #e6f4e8;
  --leaf-50:  #f3faf4;

  /* 橡实金 — 强调色 */
  --gold-700: #8a5f14;
  --gold-600: #a9781f;
  --gold-500: #c9932f;
  --gold-400: #dfae4d;
  --gold-300: #ecc87e;
  --gold-100: #fbf1d9;
  --gold-50:  #fefaf0;

  /* 中性 */
  --ink-900: #14110c;
  --ink-800: #2a251d;
  --ink-700: #4a4234;
  --ink-600: #6b6252;
  --ink-500: #8b8271;
  --ink-400: #b0a795;
  --ink-300: #d3ccbc;
  --line:    #e8e0d0;
  --line-2:  #d8cdb8;
  --bg:      #ffffff;
  --bg-soft: #faf7f1;
  --bg-mute: #f2ece1;

  --font-sans: "Inter", -apple-system, BlinkMacSystemFont, "Segoe UI",
               "PingFang SC", "Hiragino Sans GB", "Microsoft YaHei",
               "Helvetica Neue", Arial, sans-serif;
  --font-mono: "SFMono-Regular", "JetBrains Mono", Consolas, "Liberation Mono",
               Menlo, monospace;

  --radius-sm: 6px;
  --radius:    10px;
  --radius-lg: 16px;
  --radius-xl: 22px;

  --shadow-xs: 0 1px 2px rgba(43, 29, 14, .06);
  --shadow-sm: 0 2px 8px rgba(43, 29, 14, .07);
  --shadow:    0 8px 26px rgba(43, 29, 14, .10);
  --shadow-lg: 0 18px 50px rgba(43, 29, 14, .16);

  --header-h: 64px;
}

* { box-sizing: border-box; }
html { scroll-behavior: smooth; }
body {
  margin: 0;
  font-family: var(--font-sans);
  font-size: 15px;
  line-height: 1.65;
  color: var(--ink-800);
  background: var(--bg);
  -webkit-font-smoothing: antialiased;
}
a { color: var(--oak-700); text-decoration: none; }
a:hover { color: var(--oak-600); }
img { max-width: 100%; }
code, .mono { font-family: var(--font-mono); font-size: .92em; }

/* ---------- 布局 ---------- */
.container { width: 100%; max-width: 1240px; margin: 0 auto; padding: 0 24px; }
.container-narrow { max-width: 960px; }
.section { padding: 64px 0; }
.section-sm { padding: 40px 0; }
.section-soft { background: var(--bg-soft); }
.grid { display: grid; gap: 20px; }
.grid-2 { grid-template-columns: repeat(2, 1fr); }
.grid-3 { grid-template-columns: repeat(3, 1fr); }
.grid-4 { grid-template-columns: repeat(4, 1fr); }
.flex { display: flex; }
.flex-between { display: flex; justify-content: space-between; align-items: center; }
.flex-wrap { display: flex; flex-wrap: wrap; gap: 10px; }
.gap-8 { gap: 8px; }
.mt-8 { margin-top: 8px; } .mt-16 { margin-top: 16px; }
.mt-24 { margin-top: 24px; } .mt-40 { margin-top: 40px; }
.mb-8 { margin-bottom: 8px; }
.text-center { text-align: center; }
.text-sm { font-size: 13px; }
.muted { color: var(--ink-500, #6b6455); }
.small { font-size: 13px; line-height: 1.65; }
.pt-0 { padding-top: 0; }
.row-organelle td { background: var(--leaf-50, #f2f7ec); }
.tag-organelle {
  display: inline-block; margin-left: 6px; padding: 1px 7px; border-radius: 999px;
  font-size: 11px; font-weight: 500; letter-spacing: .02em;
  background: var(--leaf-100, #dcebc9); color: var(--leaf-800, #3b5a1e);
}
.text-muted { color: var(--ink-500); }
.fw-600 { font-weight: 600; }

/* ---------- 顶部导航 ---------- */
.qro-header {
  position: sticky; top: 0; z-index: 900;
  background: rgba(255, 255, 255, .92);
  backdrop-filter: saturate(180%) blur(12px);
  border-bottom: 1px solid var(--line);
}
.qro-header .container {
  height: var(--header-h);
  display: flex; align-items: center; gap: 28px;
}
.brand { display: flex; align-items: center; gap: 10px; flex: 0 0 auto; }
.brand-mark {
  width: 34px; height: 34px; border-radius: 9px; flex: 0 0 34px;
  background: linear-gradient(135deg, var(--oak-700), var(--leaf-600));
  display: flex; align-items: center; justify-content: center;
}
.brand-mark svg { width: 20px; height: 20px; }
.brand-name { font-weight: 800; font-size: 16.5px; color: var(--ink-900); line-height: 1.1; }
.brand-sub { font-size: 11px; color: var(--ink-500); line-height: 1.2; }
.qro-nav { display: flex; align-items: center; gap: 4px; margin-left: auto; }
.qro-nav a {
  padding: 7px 12px; border-radius: 8px; font-size: 14px; font-weight: 500;
  color: var(--ink-700); white-space: nowrap;
}
.qro-nav a:hover { background: var(--oak-50); color: var(--oak-700); }
.qro-nav a.active { background: var(--oak-100); color: var(--oak-800); font-weight: 600; }
.nav-toggle { display: none; margin-left: auto; background: none; border: 0; font-size: 22px; cursor: pointer; color: var(--ink-700); }

/* ---------- Hero ---------- */
.hero {
  position: relative; overflow: hidden;
  padding: 68px 0 56px;
  background:
    radial-gradient(900px 380px at 12% -8%, rgba(201,147,47,.16), transparent 62%),
    radial-gradient(760px 340px at 92% 4%, rgba(47,107,58,.14), transparent 60%),
    linear-gradient(180deg, var(--oak-50), #fff 72%);
  border-bottom: 1px solid var(--line);
}
.hero-inner { max-width: 780px; }
.hero-tag {
  display: inline-flex; align-items: center; gap: 9px; flex-wrap: wrap;
  font-size: 12.5px; color: var(--ink-600); margin-bottom: 16px;
}
.hero h1 {
  margin: 0 0 14px; font-size: clamp(30px, 4.4vw, 46px); line-height: 1.14;
  letter-spacing: -.02em; font-weight: 800; color: var(--ink-900);
}
.hero .grad {
  background: linear-gradient(100deg, var(--oak-700), var(--leaf-600) 58%, var(--gold-600));
  -webkit-background-clip: text; background-clip: text; color: transparent;
}
.hero-sub { font-size: 16px; color: var(--ink-600); max-width: 680px; margin: 0 0 10px; }
.hero-latin { font-size: 13px; color: var(--ink-500); font-style: italic; margin: 0 0 26px; }

.hero-search {
  display: flex; gap: 8px; align-items: stretch; flex-wrap: wrap;
  background: #fff; padding: 8px; border-radius: var(--radius-lg);
  box-shadow: var(--shadow); border: 1px solid var(--line);
}
.hero-search input[type=text] {
  flex: 1 1 260px; min-width: 180px; border: 0; outline: none;
  font: inherit; font-size: 15px; padding: 10px 12px; color: var(--ink-900);
  background: transparent;
}
.hero-search select, .select {
  border: 1px solid var(--line-2); border-radius: var(--radius-sm);
  padding: 8px 10px; font: inherit; font-size: 13.5px; background: #fff; color: var(--ink-800);
}
.search-suggest { display: flex; align-items: center; gap: 8px; flex-wrap: wrap; margin-top: 14px; font-size: 12.5px; color: var(--ink-500); }
.search-suggest button {
  border: 1px dashed var(--line-2); background: #fff; color: var(--oak-700);
  border-radius: 20px; padding: 3px 11px; font-size: 12px; cursor: pointer; font-family: var(--font-mono);
}
.search-suggest button:hover { border-style: solid; background: var(--oak-50); }

/* ---------- 导出框（下载页，复用首页搜索框样式） ---------- */
.export-block + .export-block { margin-top: 24px; padding-top: 22px; border-top: 1px dashed var(--line-2); }
.export-label { font-size: 13.5px; font-weight: 600; color: var(--ink-700); margin-bottom: 9px; }
.export-block .hero-search { box-shadow: var(--shadow-sm); border-radius: var(--radius); padding: 6px; }
.export-block .search-suggest { margin-top: 10px; }

/* ---------- 按钮 / 徽标 ---------- */
.btn {
  display: inline-flex; align-items: center; justify-content: center; gap: 7px;
  border: 1px solid transparent; border-radius: var(--radius);
  padding: 10px 18px; font: inherit; font-size: 14px; font-weight: 600;
  cursor: pointer; transition: all .16s ease; white-space: nowrap;
}
.btn-primary { background: var(--oak-700); color: #fff; }
.btn-primary:hover { background: var(--oak-800); color: #fff; }
.btn-leaf { background: var(--leaf-600); color: #fff; }
.btn-leaf:hover { background: var(--leaf-700); color: #fff; }
.btn-outline { background: #fff; border-color: var(--line-2); color: var(--oak-700); }
.btn-outline:hover { border-color: var(--oak-500); background: var(--oak-50); }
.btn-sm { padding: 6px 12px; font-size: 13px; }
.btn-lg { padding: 13px 26px; font-size: 15px; }
.btn-block { display: flex; width: 100%; }

.badge {
  display: inline-flex; align-items: center; gap: 5px;
  background: var(--leaf-100); color: var(--leaf-700);
  border-radius: 20px; padding: 2px 10px; font-size: 11.5px; font-weight: 600;
}
.badge-gold { background: var(--gold-100); color: var(--gold-700); }
.badge-oak { background: var(--oak-100); color: var(--oak-800); }
.badge-mute { background: var(--bg-mute); color: var(--ink-600); }
.badge-outline { background: #fff; border: 1px solid var(--line-2); color: var(--ink-600); }

/* ---------- 统计条 ---------- */
.stats { padding: 30px 0 8px; background: #fff; }
.stats-grid { display: grid; grid-template-columns: repeat(4, 1fr); gap: 18px; }
.stat-item {
  background: linear-gradient(180deg, #fff, var(--oak-50));
  border: 1px solid var(--line); border-radius: var(--radius-lg);
  padding: 18px 20px; text-align: center;
}
.stat-num { font-size: 30px; font-weight: 800; color: var(--oak-700); line-height: 1.1; letter-spacing: -.02em; }
.stat-num .unit { font-size: 15px; font-weight: 600; color: var(--ink-500); margin-left: 2px; }
.stat-label { font-size: 12.5px; color: var(--ink-500); margin-top: 4px; }

/* ---------- 区块标题 ---------- */
.sec-head { margin-bottom: 30px; }
.sec-head.center { text-align: center; max-width: 760px; margin-left: auto; margin-right: auto; }
.sec-eyebrow {
  font-size: 11.5px; font-weight: 700; letter-spacing: .12em; text-transform: uppercase;
  color: var(--gold-600); margin-bottom: 8px;
}
.sec-title { margin: 0 0 10px; font-size: 28px; font-weight: 800; color: var(--ink-900); letter-spacing: -.015em; line-height: 1.25; }
.sec-desc { margin: 0; color: var(--ink-600); font-size: 15px; }

/* ---------- 模块卡 ---------- */
.modules { display: grid; grid-template-columns: repeat(auto-fit, minmax(250px, 1fr)); gap: 18px; }
.mod-card {
  display: block; background: #fff; border: 1px solid var(--line);
  border-radius: var(--radius-lg); padding: 22px; transition: all .18s ease;
}
.mod-card:hover { border-color: var(--oak-400); box-shadow: var(--shadow); transform: translateY(-2px); }
.mod-ico {
  width: 44px; height: 44px; border-radius: 12px; margin-bottom: 14px;
  background: var(--leaf-50); color: var(--leaf-600);
  display: flex; align-items: center; justify-content: center;
}
.mod-ico svg { width: 22px; height: 22px; }
.mod-title { font-size: 16.5px; font-weight: 700; color: var(--ink-900); }
.mod-en { font-size: 11.5px; color: var(--ink-400); letter-spacing: .08em; text-transform: uppercase; margin-bottom: 8px; }
.mod-desc { font-size: 13.4px; color: var(--ink-600); min-height: 58px; }
.mod-meta { display: flex; flex-wrap: wrap; gap: 6px; margin-top: 12px; }
.mod-meta span { background: var(--oak-50); color: var(--oak-700); border-radius: 6px; padding: 2px 8px; font-size: 11.5px; }

/* ---------- 页面头（内页） ---------- */
.page-hero {
  padding: 40px 0 26px; border-bottom: 1px solid var(--line);
  background: linear-gradient(180deg, var(--oak-50), #fff);
}
.page-title { margin: 0 0 6px; font-size: 30px; font-weight: 800; color: var(--ink-900); letter-spacing: -.015em; }
.page-en { font-size: 12px; letter-spacing: .1em; text-transform: uppercase; color: var(--gold-600); font-weight: 700; margin-bottom: 8px; }
.page-desc { margin: 0; color: var(--ink-600); max-width: 820px; font-size: 14.6px; }
.page-stats { display: flex; flex-wrap: wrap; gap: 26px; margin-top: 20px; }
.page-stat .v { font-size: 22px; font-weight: 800; color: var(--oak-700); line-height: 1.1; }
.page-stat .k { font-size: 12px; color: var(--ink-500); }

/* ---------- 面板 / 表格 ---------- */
.panel {
  background: #fff; border: 1px solid var(--line); border-radius: var(--radius-lg);
  overflow: hidden; margin-bottom: 20px;
}
.panel-head {
  display: flex; align-items: center; gap: 10px; flex-wrap: wrap;
  padding: 14px 18px; border-bottom: 1px solid var(--line); background: var(--bg-soft);
}
.panel-title { font-size: 15.5px; font-weight: 700; color: var(--ink-900); display: flex; align-items: center; gap: 8px; }
.pt-bar { width: 3px; height: 15px; border-radius: 2px; background: var(--oak-600); display: inline-block; }
.panel-sub { font-size: 12.5px; color: var(--ink-500); margin-left: auto; }
.panel-body { padding: 18px; }

.data-table { width: 100%; border-collapse: collapse; font-size: 13.6px; }
.data-table th, .data-table td {
  padding: 9px 12px; border-bottom: 1px solid var(--line); text-align: left; vertical-align: top;
}
.data-table thead th {
  background: var(--bg-soft); font-weight: 700; color: var(--ink-700);
  position: sticky; top: 0; font-size: 12.6px;
}
.data-table tbody tr:hover { background: var(--oak-50); }
.data-table td.num, .data-table th.num { text-align: right; font-variant-numeric: tabular-nums; }
.table-wrap { max-height: 560px; overflow: auto; }
.table-scroll { overflow-x: auto; }

.kv { display: grid; grid-template-columns: 150px 1fr; gap: 8px 16px; font-size: 14px; }
.kv dt { color: var(--ink-500); font-weight: 600; }
.kv dd { margin: 0; color: var(--ink-800); word-break: break-word; }

/* ---------- 基因结构图 ---------- */
.gene-track { margin-top: 10px; }
.gene-track-row { display: flex; align-items: stretch; gap: 12px; margin-bottom: 10px; }
.gene-track-label {
  flex: 0 0 190px; font-size: 12.4px; color: var(--ink-600);
  font-family: var(--font-mono); word-break: break-all; padding-top: 2px;
}
.gene-track-canvas { flex: 1; position: relative; height: 34px; background: var(--bg-mute); border-radius: 5px; }
.gene-track-line { position: absolute; top: 50%; left: 0; right: 0; height: 1px; background: var(--line-2); }
.gene-exon { position: absolute; top: 7px; height: 20px; border-radius: 3px; background: var(--leaf-500); }
.gene-cds { position: absolute; top: 4px; height: 26px; border-radius: 3px; background: var(--oak-600); }
.gene-utr { position: absolute; top: 11px; height: 12px; border-radius: 3px; background: var(--leaf-300); }
.gene-ruler { position: relative; height: 22px; font-size: 10.5px; color: var(--ink-400); }
.gene-ruler span { position: absolute; transform: translateX(-50%); white-space: nowrap; }

/* ---------- JBrowse 容器 ---------- */
.jb-frame { width: 100%; height: 720px; border: 1px solid var(--line); border-radius: var(--radius-lg); background: #fff; }
.jb-bar { display: flex; align-items: center; gap: 10px; flex-wrap: wrap; margin-bottom: 12px; }
.jb-bar input[type=text] {
  flex: 1 1 240px; border: 1px solid var(--line-2); border-radius: var(--radius-sm);
  padding: 8px 11px; font: inherit; font-size: 13.5px; font-family: var(--font-mono);
}

/* ---------- 时间线 ---------- */
.timeline { display: flex; flex-direction: column; gap: 14px; position: relative; padding-left: 18px; }
.timeline::before { content: ""; position: absolute; left: 4px; top: 6px; bottom: 6px; width: 2px; background: var(--line); }
.tl-item { position: relative; }
.tl-item::before {
  content: ""; position: absolute; left: -18px; top: 6px; width: 10px; height: 10px;
  border-radius: 50%; background: #fff; border: 2px solid var(--oak-500);
}
.tl-date { font-size: 11.5px; color: var(--gold-600); font-weight: 700; }
.tl-title { font-size: 14px; font-weight: 600; color: var(--ink-900); }
.tl-desc { font-size: 13px; color: var(--ink-600); }

/* ---------- CTA / 页脚 ---------- */
.cta {
  background: linear-gradient(120deg, var(--oak-800), var(--leaf-700));
  border-radius: var(--radius-xl); padding: 38px 40px; color: #fff;
  display: flex; align-items: center; gap: 24px; flex-wrap: wrap; justify-content: space-between;
}
.cta h2 { margin: 0 0 6px; font-size: 24px; font-weight: 800; }
.cta p { margin: 0; opacity: .88; font-size: 14px; max-width: 560px; }
.cta-actions { display: flex; gap: 10px; flex-wrap: wrap; }

.qro-footer { background: var(--ink-900); color: #cfc7b8; padding: 44px 0 22px; margin-top: 40px; }
.qro-footer a { color: #e3dccd; }
.qro-footer a:hover { color: #fff; }
.footer-top { display: grid; grid-template-columns: 1.6fr 1fr 1fr 1fr; gap: 30px; padding-bottom: 26px; border-bottom: 1px solid rgba(255,255,255,.10); }
.footer-brand .brand-name { color: #fff; }
.footer-col h4 { font-size: 13px; color: #fff; margin: 0 0 10px; font-weight: 700; }
.footer-col ul { list-style: none; margin: 0; padding: 0; display: flex; flex-direction: column; gap: 6px; font-size: 13px; }
.footer-bottom { display: flex; justify-content: space-between; gap: 16px; flex-wrap: wrap; padding-top: 18px; font-size: 12px; color: #9b9484; }

/* ---------- 空状态 / 提示 ---------- */
.empty { padding: 36px; text-align: center; color: var(--ink-500); font-size: 14px; }
.notice {
  background: var(--gold-50); border: 1px solid var(--gold-300); color: var(--gold-700);
  border-radius: var(--radius); padding: 10px 14px; font-size: 13.4px;
}

/* ---------- 响应式 ---------- */
@media (max-width: 1000px) {
  .grid-4 { grid-template-columns: repeat(2, 1fr); }
  .grid-3 { grid-template-columns: repeat(2, 1fr); }
  .stats-grid { grid-template-columns: repeat(2, 1fr); }
  .footer-top { grid-template-columns: 1fr 1fr; }
}
@media (max-width: 760px) {
  .qro-nav { display: none; }
  .qro-nav.open {
    display: flex; flex-direction: column; align-items: stretch;
    position: absolute; top: var(--header-h); left: 0; right: 0;
    background: #fff; border-bottom: 1px solid var(--line); padding: 10px 24px 16px;
    box-shadow: var(--shadow);
  }
  .nav-toggle { display: block; }
  .grid-2, .grid-3, .grid-4 { grid-template-columns: 1fr; }
  .stats-grid { grid-template-columns: repeat(2, 1fr); }
  .hero { padding: 44px 0 34px; }
  .kv { grid-template-columns: 1fr; }
  .gene-track-label { flex: 0 0 110px; font-size: 11px; }
  .jb-frame { height: 560px; }
}